''Pterocaesio marri'', Marr's fusilier, bigtail fusilier, blacktip fusilier, bananafish or twinstripe fusilier is a species of marine

ray-finned fish Actinopterygii (; ), members of which are known as ray-finned fishes, is a class of bony fish. They comprise over 50% of living vertebrate species. The ray-finned fishes are so called because their fins are webs of skin supported by bony or hor ...

, a fusilier belonging to the

family Family (from la, familia) is a Social group, group of people related either by consanguinity (by recognized birth) or Affinity (law), affinity (by marriage or other relationship). The purpose of the family is to maintain the well-being of its ...

Caesionidae Caesionidae, the fusiliers, are a family of marine ray-finned fishes in the order Perciformes. The family includes about 23 species. They are related to the snappers, but adapted for feeding on plankton, rather than on larger prey. They are fou ...

. It is widespread around reefs in the

Indo-West Pacific The Indo-Pacific is a vast biogeographic region of Earth. In a narrow sense, sometimes known as the Indo-West Pacific or Indo-Pacific Asia, it comprises the tropical waters of the Indian Ocean, the western and central Pacific Ocean, and the ...

region.

Taxonomy

''Pterocaesio marri'' was first formally described in 1953 by the

American American(s) may refer to: * American, something of, from, or related to the United States of America, commonly known as the "United States" or "America" ** Americans, citizens and nationals of the United States of America ** American ancestry, pe ...

ichthyologist Ichthyology is the branch of zoology devoted to the study of fish, including bony fish ( Osteichthyes), cartilaginous fish (Chondrichthyes), and jawless fish (Agnatha). According to FishBase, 33,400 species of fish had been described as of Octobe ...

Leonard Peter Schultz Leonard Peter Schultz (1901–1986) was an American ichthyologist. Biography Schultz was born in 1901, at Albion, Michigan. He received education on ichthyology at Albion College, in which he got his bachelor's degree, in 1924. In 1926, he got ...

with the type localities given as a lagoon a quarter mile off Amen Island in

Bikini Atoll Bikini Atoll ( or ; Marshallese: , , meaning "coconut place"), sometimes known as Eschscholtz Atoll between the 1800s and 1946 is a coral reef in the Marshall Islands consisting of 23 islands surrounding a central lagoon. After the Second ...

in the

Marshall Islands The Marshall Islands ( mh, Ṃajeḷ), officially the Republic of the Marshall Islands ( mh, Aolepān Aorōkin Ṃajeḷ),'' () is an independent island country and microstate near the Equator in the Pacific Ocean, slightly west of the Internati ...

. In his 1987 review of the Caesionidae, Kent E. Carpenter placed this species within the

subgenus In biology, a subgenus (plural: subgenera) is a taxonomic rank directly below genus. In the International Code of Zoological Nomenclature, a subgeneric name can be used independently or included in a species name, in parentheses, placed between t ...

''Squamosicaesio'', of which it is the

type species In zoological nomenclature, a type species (''species typica'') is the species name with which the name of a genus or subgenus is considered to be permanently taxonomically associated, i.e., the species that contains the biological type specimen ...

. The specific name honours John C. Marr of the

U.S. Fish and Wildlife Service The United States Fish and Wildlife Service (USFWS or FWS) is an agency within the United States Department of the Interior dedicated to the management of fish, wildlife, and natural habitats. The mission of the agency is "working with othe ...

, who had oversight of the commercial fisheries at Bikini Atoll.

Description

''Pterocaesio marri'' has a

fusiform Fusiform means having a spindle-like shape that is wide in the middle and tapers at both ends. It is similar to the lemon-shape, but often implies a focal broadening of a structure that continues from one or both ends, such as an aneurysm on a b ...

and elongated body which is moderately laterally compressed. There are small conical teeth in the jaws but none on the vomer and palatines. The

dorsal fin A dorsal fin is a fin located on the back of most marine and freshwater vertebrates within various taxa of the animal kingdom. Many species of animals possessing dorsal fins are not particularly closely related to each other, though through conv ...

contains 10-11 spines and 14-16 soft rays while the

anal fin Fins are distinctive anatomical features composed of bony spines or rays protruding from the body of a fish. They are covered with skin and joined together either in a webbed fashion, as seen in most bony fish, or similar to a flipper, as se ...

has 3 spines and 11-13 soft rays. There are scales on both the dorsal and anal fins. There are 2-24 rays in the

pectoral fin Fins are distinctive anatomical features composed of bony spines or rays protruding from the body of a fish. They are covered with skin and joined together either in a webbed fashion, as seen in most bony fish, or similar to a flipper, as ...

s. This species attains a maximum

total length Fish measurement is the measuring of individual fish and various parts of their anatomies. These data are used in many areas of ichthyology, including taxonomy and fisheries biology. Overall length * Standard length (SL) is the length of a fish ...

of . The overall colour is blue with dark tips to the lobes of the

caudal fin Fins are distinctive anatomical features composed of bony spines or rays protruding from the body of a fish. They are covered with skin and joined together either in a webbed fashion, as seen in most bony fish, or similar to a flipper, as se ...

and two thin yellow to brownish stripes along the flanks, the lowest stripe largely running along the

lateral line The lateral line, also called the lateral line organ (LLO), is a system of sensory organs found in fish, used to detect movement, vibration, and pressure gradients in the surrounding water. The sensory ability is achieved via modified epithelial ...

Distribution and habitat

''Pterocaesio marri'' has a wide range in the

. It is occurs along the coast of East Africa from Somalia south to South Africa, although they are absent from the Red Sea and the Persian Gulf. They are found east across the Indian Ocean into the Pacific Ocean as far east as the

Marquesas Islands The Marquesas Islands (; french: Îles Marquises or ' or '; Marquesan: ' ( North Marquesan) and ' ( South Marquesan), both meaning "the land of men") are a group of volcanic islands in French Polynesia, an overseas collectivity of France in th ...

. It has also been recorded from

New Caledonia ) , anthem = "" , image_map = New Caledonia on the globe (small islands magnified) (Polynesia centered).svg , map_alt = Location of New Caledonia , map_caption = Location of New Caledonia , mapsize = 290px , subdivision_type = Sovereign st ...

, Fiji and Tonga and south to the northern

Great Barrier Reef The Great Barrier Reef is the world's largest coral reef system composed of over 2,900 individual reefs and 900 islands stretching for over over an area of approximately . The reef is located in the Coral Sea, off the coast of Queensland, ...

Queensland ) , nickname = Sunshine State , image_map = Queensland in Australia.svg , map_caption = Location of Queensland in Australia , subdivision_type = Country , subdivision_name = Australia , established_title = Before federation , established_ ...

and the reefs in the

Coral Sea The Coral Sea () is a marginal sea of the South Pacific off the northeast coast of Australia, and classified as an interim Australian bioregion. The Coral Sea extends down the Australian northeast coast. Most of it is protected by the Fre ...

, as well as

Christmas Island Christmas Island, officially the Territory of Christmas Island, is an Australian external territory comprising the island of the same name. It is located in the Indian Ocean, around south of Java and Sumatra and around north-west of the ...

. This species occars at depths between on reefs and around oceanic islands.

Biology

''Pterocaesio marri'' forms

schools A school is an educational institution designed to provide learning spaces and learning environments for the teaching of students under the direction of teachers. Most countries have systems of formal education, which is sometimes compulsor ...

in midwater which forage for

zooplankton Zooplankton are the animal component of the planktonic community ("zoo" comes from the Greek word for ''animal''). Plankton are aquatic organisms that are unable to swim effectively against currents, and consequently drift or are carried along by ...

. They are

oviparous Oviparous animals are animals that lay their eggs, with little or no other embryonic development within the mother. This is the reproductive method of most fish, amphibians, most reptiles, and all pterosaurs, dinosaurs (including birds), and ...

laying large numbers of small pelagic eggs.

Fisheries

''Pterocaesio marri'' is subject to heavy fishing pressures in parts of its range, such as the Philippines. Fishers use

drive-in net A fishing net is a net used for fishing. Nets are devices made from fibers woven in a grid-like structure. Some fishing nets are also called fish traps, for example fyke nets. Fishing nets are usually meshes formed by knotting a relatively thin ...

s in western

Palawan Palawan (), officially the Province of Palawan ( cyo, Probinsya i'ang Palawan; tl, Lalawigan ng Palawan), is an archipelagic province of the Philippines that is located in the region of Mimaropa. It is the largest province in the country in ...

but it is also taken using

gill net Gillnetting is a fishing method that uses gillnets: vertical panels of netting that hang from a line with regularly spaced floaters that hold the line on the surface of the water. The floats are sometimes called "corks" and the line with corks is ...

s and

fish trap A fish trap is a trap used for fishing. Fish traps include fishing weirs, lobster traps, and some fishing nets such as fyke nets. Traps are culturally almost universal and seem to have been independently invented many times. There are two main ...

s in other parts of its range. It is sometimes used by the Indian Ocean and West Pacific tuna fisheries as bait.

As food

''Pterocaesio marri'' is often served as

fried fish Fried fish is any fish or shellfish that has been prepared by frying. Often, the fish is covered in batter, egg and breadcrumbs, flour, or herbs and spices before being fried and served, often with a slice of lemon. Fish is fried in many part ...

Okinawa Prefecture is a prefecture of Japan. Okinawa Prefecture is the southernmost and westernmost prefecture of Japan, has a population of 1,457,162 (as of 2 February 2020) and a geographic area of 2,281 km2 (880 sq mi). Naha is the capital and largest city o ...

Japan Japan ( ja, 日本, or , and formally , ''Nihonkoku'') is an island country in East Asia. It is situated in the northwest Pacific Ocean, and is bordered on the west by the Sea of Japan, while extending from the Sea of Okhotsk in the north ...

References

External links

Photos of ''Pterocaesio marri'' on FishBase
{{Taxonbar, from=Q2265577 marri Fish described in 1953